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Tree Biology and Pruning Basics
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Page 1
Tree Biology and
Pruning Basics
James Luce
City of Bellingham
Parks Arborist
Page 2
Tree Biology
References:
Dr. Ed Gilman and Scott Jones, University of Florida
Paul Reis, Urban Forester, Oregon Dept. of Forestry
Dr. Alex Shigo, author of A New Tree Biology
Page 3
Forest conditions are optimum for trees
Page 4
However,
properly
maintained trees
add value to your
landscape
Streets and
urban conditions
are far from
optimum!
Page 6
Decision making must be based on knowledge and experience.
Page 7
What is tree biology?
The study of the Life Processes
of a tree.
Including a study of the GROWTH,
STRUCTURE, EVOLUTION, etc. of a tree.
Page 8
Leaves work like solar panels.
They “feed” the tree by converting sunlight into energy; glucose.
Page 9
• Trees make their food with the energy of
sunlight; a process known as photosynthesis.
• Nutrients and water are taken through the roots.
• Carbon is taken from the CO2 in the air.
Page 10
Life processes – some examples:
photosynthesis –
a tree’s gotta eat!
support
mycorrhizal interactions
Page 11
The tree below the ground: Roots support and adsorb water and nutrients.
Page 12
Soil
• Soil is the matrix to support the roots and the community of
• organisms that are part of a healthy tree.
• All trees need mycorrhizal associations – healthy soils have a huge
diversity of organisms including fungi, bacteria
• Must have air spaces and the ability to drain yet hold moisture.
• Often the biggest challenge in urban landscapes.
• Organic matter is the food that feeds the soil food web.
Page 14
Mycorrhizae
are organs
that function
in
synergistic
ways.
In other words, they work
with the roots.
Page 15
Mycorrhizal fungi assist tree roots in
gathering (adsorb) nutrients & are thought to
help protect the tree.
Page 16
Thread-like mycorrhyzae hyphae multiple the adsorptive
area by a factor of thousands!
Page 17
Morels, chanterelles, truffles are fruiting
bodies of mycorrhyzal fungi.
Page 19
Mycorrhizae
do not
fix nitrogen
Page 20
Actinorhizal plants, alders and legume family trees, do fix nitrogen (makes
nitrogen available in the soil) and are great pioneer species.
Page 21
A new tree grows around the old tree with every cycle. Length and height are added and then augmented.
Page 22
Secondary Growth Anatomy &
Tree Rings
By Kim D. Coder, Professor, Silvics/Ecology
Warnell School of Forest Resources
The University of Georgia August, 1999
Page 23
The timing of branch and trunk growth creates a strong interlocking
structure in branch attachments.
from A New Tree Biology Dr. Alex Shigo
Page 24
The cambium:
The cambium is the
primary meristem
producing radial
growth.
It forms the
phloem & xylem.
Page 25
The Bark:
The bark is
everything outside
the vascular
cambium.
As you can see,
there is a lot going
on in the bark.
Page 26
The Xylem (wood):
The xylem includes
everything inside the
vascular cambium.
Page 27
The Xylem: a growth increment (ring):
The rings seen in
many trees
represent one
growth increment.
Growth rings
provide the texture
seen in wood.
Page 28
Starch reserves near buds.
Page 29
Roots are not
stems.
Note the
differences.
Page 30
Leaves feed the part of the tree nearest them.
Page 31
Phenology
Timing of life processes
• Seasonal growth generally starts at the buds . The tree
parts below follow.
• Stored energy reserves (starches) are used up to push
the new growth.
• Roots can grow anytime the soil temps are above 40 F.
• Genetics and climate influence the phenology.
• Auxins are compounds made by the plants that
coordinate the growth of different parts of the plant.
• Think about the timing of growth, when making pruning
decisions
Page 32
Tree Growth
• Genetics determine basic form and size of a tree.
• Trees grow towards the sunlight and away from the
pull of gravity.
• Roots grow where they can find air and water.
• Trees grow to maintain the dynamic balance of the
tree.
• Trees grow new wood around wounds (they don’t
“heal.”
Page 33
The basic plant form is
dictated by genetics.
The conical form of this
evergreen shows strong
apical dominance –
“excurrent” growth.
The auxins from the top
suppress the growth lower
down the trunk.
If the top is removed, the
lower branches will no
longer be suppressed.
Later a new top forms
sending the chemical
message that suppresses
the lower growth.
Page 34
Apical dominance is exerted the much
the same way on later branches.
The branch tip is dominant and
suppresses the growth behind it.
If removed latent buds will sprout. The
remaining growth will grow longer.
Page 35
Decurrant form, open and spreading, on an American elm
Page 36
Why prune trees?
• Prune young trees to establish a sound structure.
– Establish a strong scaffolding, remove weak or
rubbing branches
• For tree health might remove dead and diseased
branches.
• For values other than tree health.
– Clearance for structures, aesthetics, fruit or flower
production
Page 37
A “doughnut maple” pruned for clearance.
Page 38
Trees may be pruned to promote showy flowers or better fruit yield
Page 39
Street trees grow in extreme conditions. Early pruning helps avoid future conflicts.
Page 40
Topping is not proper pruning!!
It damages trees and destroys
canopy integrity.
Page 41
How should trees be pruned?
• With forethought
• With foresight
• According to standards
• With the proper tools
• At the proper time
• With a clearly defined
purpost
Page 42
Not pruning is an option
• In native forest remnant stands, there may
not be as much of a need to prune
• Self pruning will occur
• Tree species and growth habits dictate
pruning needs
• Not pruning can cause structural problems
to go undetected
Page 43
Training for Structure and Form
• Directs growth to
fulfill landscape
function
• Reduces structural
defects that may lead
to failure
• Decreases hazard
potential and liability
risks
Page 44
Five Steps for Training Young Trees
Remove broken, dead, dying, diseased,
or damaged branches
Select and establish a central leader
Select and establish the lowest
permanent branch
Select and establish scaffold branches
Select temporary branches below the
lowest permanent branch
Page 45
The Biology Behind Tree Pruning
• Pruning is a form of wounding, thus if
done incorrectly, can be detrimental
• Correct cuts at branch collar speeds
sealing of pruning wounds
• Stubs, topping cuts, and snagged ends
are entry points for insect and disease
• Preventive pruning is less stressful for a
tree than corrective pruning
Page 46
The Biology Behind Tree Pruning
• Successful pruning wound
closure is dependent on
the quality of the cut
• The larger the pruning cut,
the larger the wound, the
longer it takes to close
Page 47
Compartmentalization
Of
Decay
In
Trees = CODIT
• Trees are highly ordered,
COMPARTMENTED
plants, that instead of
healing,
COMPARTMENTALIZE in
an orderly way the injured
and infected tissues.
From a New Tree Biology
byAlex Shigo
Page 50
Compartmentalization Styles
• Trees that grow slowly are usually good
compartmentalizers and resist decay
– Most oaks, some elms, black locust,
hornbeam, walnut, sycamore, etc
• Trees that “live fast and die young” are
usually poor compartmentalizers, and
often succumb to decay after injury.
– Birch, ash, poplars, beech, crabapples,
willows, some oaks, cherries, cottonwoods,
aspens, horse chestnut, silver maple,
redbud, etc
Page 51
Proper techniques
• Target pruning
terms:
– branch collar
– branch bark ridge
Page 52
Making proper pruning cuts
• The final cut should
be just outside the
branch collar
• Often the weight of
the branch must first
be removed with a
preliminary under-cut
Page 53
Wound closure profile
A good pruning cut will seal quickly and be shaped like a doughnut.
Page 55
In this picture, branch wood is brown. It
forms first each spring, growing on top of
last years trunk wood, yellow, which forms
later in the spring, overlapping the recently
formed branch wood.
Branch wood remains after the trunk has rotted
away, leaving. The ridges are a result of the
alternating growth of branch and trunk wood. The
ridges act like threads on a screw to secure the
branch to the trunk. These ridges only form when
the branch is much smaller than the trunk.
Page 56
Branch bark ridge
Collar: swollen area at the base of
the branch where it joins the trunk.
The tissue is rich in energy reserves
and chemicals that hinder the
spread of decay. Good pruning cuts
avoid cutting into the collar.
Collar
Illustration and Photos by Edward F. Gilman, Professor, Environmental Horticulture Department, IFAS, University of Florida
Page 57
Improper
pruning cuts
• Some pruning
cuts can cause
more damage
than if the tree
had been left
alone
– Bark Ripping
– Flush Cutting
– Stub Cutting
Page 58
Specific Hazards Related to Topping
• Trees have difficulty
in defending a large
wound
• Regrown branches
are weakly attached
• Poor structure –
topping destroys the
natural form and
taper of a tree
Page 60
Pruning Responses
When pruning, think in terms of the tree’s response to the
cuts.
• How will the tree react?
• Wound response
• Energy needed
• Where will the new growth be ?
• What will be the effect on the whole tree?
Page 62
What is good structure?
1. Forest grown vs. open grown tree
2. Codominant stems
3. Good branch attachment
4. Structurally sound tree
Outline of topics
Page 63
Trees grown in natural conditions are usually fine, without us…
Page 64
…but conditions change.
Page 65
Understanding tree structure problems
• Branch angles
• Included bark
• Co-dominant stems
Page 66
In this tree, the majority of structural branches originate at the same location. One
branch failure, and the tree will most likely be mortally wounded
Page 67
Co-dominant Stems
• Co-dominant stems,
unlike branches do not
have collars that form
protection zones.
• Cracks at co-dominant
stems are always a
sign of weakness
• Stem bark ridges are a
sign of stronger co-
dominant stems
Page 68
Included bark; a weak point.
Page 70
Bark inclusion
Decay and
discoloration
from self-
wounding and
micro-organisms
breaking down
wood.
A narrow branch angle is a perfect
environment for microorganisms; the
area stays moist, warm, and dark
Page 71
Good branch attachment
• How does a BRANCH form?
• What are the indicators?
Page 72
Collar
No branch bark ridge
Page 73
Branch bark ridge present
• Some branch
unions have
a prominent
branch bark
ridge
Page 74
Tree Structure Problems:
Included bark
• Included bark is
defined as bark that
is squeezed
between stems
• Included bark
doesn’t necessarily
need to be decayed
in order to fail
Page 75
Weak
union
Strong
union
Photos by Edward F. Gilman, Professor, Environmental Horticulture Department, IFAS, University of Florida."
Page 76
Good
structure
Page 78
Subordination
Before
pruning
After light
pruning
After moderate
pruning
The objective of structural pruning is to develop good sustainable branch and trunk form in shade trees by making
pruning cuts in the canopy at strategic and planned locations. On many trees structural pruning can be taught as a
three step process: step one – locate the stem that will make the best leader; step two – locate those stems and
branches that are competing with this chosen leader; step three – decide where to cut these competing stems
back to. Subordination or reduction of competing stems is one of the most important techniques to use to develop
good structure in shade trees. Pruning cuts are indicated as dotted lines.
Page 79
Temporary vs. permanent branch
management
(Long Term thinking)
• Keep in mind
that all low
branches will
eventually be
removed; they
are temporary.
A good pruning program will make sure that temporary branches are removed before
they grow larger than about half the trunk diameter.
Page 80
• Early canopy
training to
create a
branch
structure that
will provide
lift truck
access for
future
maintenance
Page 81
Large branch removal vs.
early structural pruning
• Leaves a huge wound
• No branch protection zone
• Access for decay
Big wounds can mean
management woes
Page 82
Over-pruning causes sprouting
• This mass of sprouts
resulted from removing
too many low branches
at one time
• Prevent this by pruning
regularly so less than
25% needs to be
removed at a time
Page 83
Lions-tailed
• Over-lifting or over-thinning leaves live branches
only at the edge of the canopy
• Tremendous numbers of sprouts often result from
this type of tree mutilation
thinning
Page 84
A few words about topping
• Structurally unsound
• Stressful to the tree
• Expensive to maintain
• Ugly
Page 85
Topping
• Not an appropriate
method of reducing
canopy size
• Uses heading cuts
through several-year-
old woody tissue
• Results in decay and
weak structure
reduction
Page 86
Topping damage
• A look inside a tree topped several years ago
reveals some of the problems with topping
Page 87
Tree Myths Undone
Trees do not heal wounds. They grow over them.
Wound dressing are not usually a good thing.
Sap bleeding from pruning cuts in the spring doesn’t hurt
the tree.
Stake new trees only as needed.
Adding fertilizer may be bad for your trees.
Not cutting down a tree doesn’t mean you haven’t killed
it. Be kind to root zones.
Page 88
THINGS THE TREES
CAN TEACH US
• TREES DO NOT MOVE AWAY FROM THEIR
PROBLEMS.
• TREES, WHEN THREATENED OR INJURED,
REPSOND RAPIDLY.
• TREES HAVE MANY FRIENDS. THEY ARE
CONNECTED IN WAYS THAT PROVIDE
SYNERGENIC ASSOCIATIONS.
• TREES LIVE WITHIN THEIR MEANS.
Page 89
For further reading
• Illustrated Guide to
Pruning by Dr. Ed Gilman
• Available from:
The Pacific NW Chapter of
the International Society of
Arboriculture, P.O. Box 811,
Silverton, OR 97381
or www.pnwisa.org
Page 90
treelink.org
treesaregood.org
Page 91
Trees
don’t
need
people.
People
need
trees.
Page 92
Thank you !
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